"The male undergoes some development while still within the mother 1. After
emergence the egg (young male) remains adherent for two or three days
to some part of the mother, fig. 7a, and finally inserts his head into his
mother's genital aperture and feeds upon here fig. 7 b. A somewhat later
stage, fig. 7 e, shows only a remnant of the mother. In the course of a
week of feeding, often less, the mother has been completely devoured
and the male is nearly ready to pupate. Such a male; fig. 8, shows clear
segmentation into head, thorax and abdomen. The length of time taken
for development in all forms, excepting the male, is a remarkably long
one. It is certainly several months at room temperature for all the female
forms and even the relatively rapid development of the male requires
from twenty to thirty days (from the shedding of the egg until eclosion)."
Scott, A.C. Paedogenesis in the coleoptera. Z. Morph. u. Okol. Tiere 33, 633–653 (1938). https://doi.org/10.1007/BF00407571
There are five reproductive types in the species described by Scott: the paedogenetic larval mother which produces a (single) male, the one which produces a brood of females, the one which produces both males and females, the metamorphosed adult male and the metamorphosed adult female. The paedogenetic forms reproduce asexually through parthenogenesis. Males are haploid and females are diploid.